Gaylord "Greg" Focker (Ben Stiller) is a nurse living in Chicago. He intends to propose to his girlfriend Pam Byrnes (Teri Polo), but his plan is disrupted when he learns that Pam's sister's fiance had asked Pam's father for permission before proposing. Greg and Pam travel to Pam's parents' house to attend Pam's sister's wedding. Greg hopes to propose to Pam in front of her family after receiving her father's permission. But this plan is put on hold when the airline loses his luggage, including the engagement ring.
At the Byrnes' home, Greg meets Pam's father Jack (Robert De Niro), mother Dina (Blythe Danner) and their beloved cat Mr. Jinx. Jack takes an instant dislike to Greg and openly criticizes him for his choice of career as a male nurse and whatever else he sees as a difference between Greg and the Byrnes family. Greg tries to impress Jack, but his efforts fail. Greg becomes even more uncomfortable after he receives an impromptu lie detector test from Jack and later learns from Pam that her father is a retired CIA counterintelligence officer.
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